Drupal Association members fund grants that make connections all over the world.
1) Configure as per screenshot
2) Add a user who can create 'article'
3) Log in as new user and create an article
4) Log in as another user and comment on newly created node
At this point I would expect user 1 to get an email saying there has been a comment on their node. It does not happen. The only way to get this to work is to log in as user 1 and edit your account and then save the preset settings. Once you do this notification works as expected.
I don't know if this is by design or mistake but it is pretty annoying having to tell every user that to get notifications they have to edit then save their profile.